Harlequins have strengthened their front row for next season with the signing of England prop Paul Doran-Jones on a two-year contract.
The 27-year-old front-rower, who has spent two seasons with Northampton, made his international debut against Argentina four years ago and has since made three further appearances for his country.
Harlequins Director of Rugby, Conor O'Shea said: "While we were disappointed to lose James Johnston we recognise the need to have an experienced player to come in to supplement the existing tight head talent we have.
"With that in mind we are very fortunate to be able to bring in a player of Paul's ability.
"He comes from a fabulous club and has been coached by one of the best forwards coaches in the country so I have no doubt he will add greatly to what we have at Harlequins."
Doran-Jones added: "I'm delighted to have the opportunity to play at the best club in the Premiership. The team play with smiles on their faces, a sure sign that they enjoy their style of rugby.
"I'm looking forward to being part of that, to improving my game and to working with the coaches and players to contribute to further success for Harlequins."